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Dipterocarpaceae
Dipterocarpus retusus
Scientific classification
Kingdom:	Plantae
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Rosids
Order:	Malvales
Family:	Dipterocarpaceae Blume[1]
Genera
Anisoptera Cotylelobium Dipterocarpus Dryobalanops Hopea Marquesia Monotes Neobalanocarpus Parashorea Pseudomonotes Shorea Stemonoporus Upuna Vateria Vateriopsis Vatica

Dipterocarpaceae are a family of 16 genera and approximately 695 known species^[2] of mainly tropical lowland rainforest trees. The family name, from the type genus Dipterocarpus, is derived from Greek (di = two, pteron = wing and karpos = fruit) and refers to the two-winged fruit. The largest genera are Shorea (196 species), Hopea (104 species), Dipterocarpus (70 species), and Vatica (65 species).^[3] Many are large forest emergent species, typically reaching heights of 40–70 m, some even over 80 m (in the genera Dryobalanops,^[3]Hopea^[4] and Shorea),^[4] with the tallest known living specimen (Shorea faguetiana) 93.0 m tall.^[4] The species of this family are of major importance in the timber trade. Their distribution is pantropical, from northern South America to Africa, the Seychelles, India, Indochina, Indonesia and Malaysia.^[5] The greatest diversity of Dipterocarpaceae occurs in Borneo.^[6] Some species are now endangered as a result of overcutting, extensive illegal logging and habitat conversion. They provide valuable woods, aromatic essential oils, balsam, resins and are a source for plywood.

Classification

The dipterocarp family is generally divided into two subfamilies:

• Dipterocarpoideae: the largest of the subfamilies, it contains 13 genera and about 475 species. Distribution includes the Seychelles, Sri Lanka, India, Southeast Asia to New Guinea, and a large distribution in Borneo, where they form the dominant species in the lowland forests. North Borneo (Brunei, Sabah and Sarawak) is the richest area in the world for dipterocarp species.^[3] The Dipterocarpoideae can be divided morphologically into two groups,^[6][7] and the tribe names Shoreae and Dipterocarpeae are sometimes used, but genetic evidence so far does not support this division:^[8][9]
  • 
    Valvate - Dipterocarpeae group (Anisoptera, Cotylelobium, Dipterocarpus, Stemonoporus, Upuna, Vateria, Vateriopsis, Vatica). The genera of this group have valvate sepals in fruit, solitary vessels, scattered resin canals, and basic chromosome number x = 11.
  
  
  • 
    Imbricate - Shoreae group (Balanocarpus, Hopea, Parashorea, Shorea). The genera of this group have imbricate sepals in fruit, grouped vessels, resin canals in tangential bands, and basic chromosome number x = 7. A recent molecular study suggest that the genus Hopea forms a clade with Shorea sections Anthoshorea and Doona, and should be merged into Shorea.^[8]
  
  


• 
  Monotoideae: 3 genera, 30 species. Marquesia is native to Africa. Monotes has 26 species, distributed across Africa and Madagascar. Pseudomonotes is native to the Colombian Amazon.


• 
  Pakaraimoideae: formerly placed here contains a single species, Pakaraimaea roraimae, found in the Guaianan highlands of South America. It is now found to be more closely related to Cistaceae and is placed there in the APG IV (2016)^[10]

A recent genetic study found that the Asian dipterocarps share a common ancestor with the Sarcolaenaceae, a tree family endemic to Madagascar.^[11] This suggests that ancestor of the Dipterocarps originated in the southern supercontinent of Gondwana, and that the common ancestor of the Asian dipterocarps and the Sarcolaenaceae was found in the India-Madagascar-Seychelles land mass millions of years ago, and were carried northward by India, which later collided with Asia and allowed the dipterocarps to spread across Southeast Asia and Malaysia. The first dipterocarp pollen has been found in Myanmar (which at that time was part of the Indian plate) and it dates from the upper Oligocene.^[12] The sample appears to slowly increase in terms of diversity and abundance across the region into the mid-Miocene^[12] Chemical traces of dipterocarp resins have been found dating back to the Eocene of India.

Fossilized arthropods

52-million-year-old amber found in the Gujarat province, India, containing a large amount of fossilized arthropods, was identified as sap from the Dipterocarpaceae family.^[13]

Ecology

Dipterocarpaceae species can be either evergreen or deciduous.^[14] Species occurring in Thailand grows from sea level to c. 1300 m elevation. Environments in which the species of the family occur in Thailand include: Lowland dipterocarp forest 0–350 m; Riparian fringe; Limestone hills; and Coastal hills.

See also

• Dipterocarp timber classification

References

External links

• 
  Media related to Dipterocarpaceae at Wikimedia Commons


• 
  Data related to Dipterocarpaceae at Wikispecies


• 
  Center for International Forestry Research (1998). A Review of Dipterocarps: Taxonomy, ecology and silviculture (PDF version) (PDF). ISBN 978-979-8764-20-2.
  


• Dipterocarpaceae Data Base